Grizzly
Bear Habitat
An
Adaptive Management Approach for Integrating Grizzly Bear
Habitat Requirements and Silvicultural Practices in Coastal
B.C.
Contacts:
Tony
Hamilton (Wildlife Branch - Victoria)
Ralph Winter - Silviculture
Hal Reveley - Silviculture Development Forester
Ted Nash - Silviculture - Vancouver F.R.
Donald S. McLennan principle worker for the project
Tom Johnson principle worker for the project
Background:
The
project was initiated in 1992 to develop silvicultural systems
that provide grizzly forage over the entire rotation of
a managed stand, while, at the same time, producing high
quality timber. As a result of the wide ecological variation
on which the reduced stocking standards can be applied,
silvicultural systems will be developed on a site- specific
basis.
Objective:
The
main objective of the project is to evaluate the effect
of varying gap size and the numbers of trees per cluster
within trial plantations on light conditions, and the species
composition and productivity of forage species within the
gaps created.
The
second primary objective is to evaluate the influence of
varying inter-cluster distances, and the numbers of trees
per cluster on both timber quantity and quality within the
plantation.
Specific
objectives of the project are:
a)
to evaluate the effect of varying gap size on;
-
i)
light levels (photosynthetically-active radiation -PAR)
within the gaps, both at the onset of the experiment,
and as the plantation develops;
-
ii)
the production, on a per hectare basis, of both the
quantity and quality of grizzly bear forage;
-
iii)
the species of forage plants that occur in the gaps,
and how this changes with time.
b)
to evaluate the influence of varying inter-cluster distances
and the numbers of trees per cluster on both timber quantity
and quality within the plantation, and, specifically, to;
-
i)
assess the effects of different numbers of trees per
cluster;
-
ii)
assess the effects of different species mixtures within
clusters; and,
-
iii)
assess the effects of different pruning strategies.
Specific
Objectives for 1993-1994 were;
-
1.
to document changes in light conditions and the response
of forage species in naturally-occurring gaps in unmanaged
stands, along a chronosequence of stands in 2 site associations.
Experimental
Design:
In
the context of this study, the management objective is to
regenerate forest stands that have forage value over the
entire rotation, while still producing high-quality timber.
There is uncertainty associated with achieving this objective
because a number of important factors that will determine
the outcome are unknown. For example, it is not known how
the proposed stocking levels will affect light conditions,
and how these changes in light levels will affect the productivity
of the different forage species. It is also not known how
reduced stocking, whether clustered or widely-spaced, will
affect timber quantity and quality, and how much stand tending
will be required.
Given
these uncertainties, the adaptive management approach to
be taken is to initially, set up relatively small trials
(4-10 ha) at a number of different spacing and cluster arrangements,
and on a range of site associations. These initial blocks
will be established at formal, informal and monitoring levels,
and information will be gathered at intervals as the plantations
are monitored. The trials will be monitored for forage production
and growth and yield of crop trees to begin to identify
the most promising methods. At the same time it is proposed
that retrospective research be carried out to provide additional
information from older, naturally-regenerated stands.
Conclusions
based on this data will be used to alter or fine-tune the
development of the most appropriate silvicultural systems.
When original levels of uncertainty are reduced, then larger
operational trials will be established with fewer treatments.
It is expected that the overall objectives can only be achieved
by monitoring a number of experimental sites over a complete
rotation. Eventually, the most promising approaches for
each of the site associations will be developed and become
the operational standard.
The
Working Plan outlines the approach to be taken, and describes
the layout and experimental design of the trials. Please
refer to this (McLennan and Johnson, 1993) document for
more detailed information.
The
generalized null hypotheses to be tested in the proposed
trials are;
Ho1: that forage values in treatments and control blocks
are the same
Ho2: that crop tree growth performance in treatments
and control blocks are the same
As
discussed in the Working Plan, replication for the formal
trials will occur within site associations over a number
of years as blocks become available. This distribution means
that, in any given year of measurement, trials will be of
different age, and will not be directly replicated within
the same trial area. Because of this it is proposed that
non- parametric statistical tests, such as the Kruskall-Wallis
test for k>2 treatments, or the Wilcoxin-Mann-Whitney
two-sample test, be used to compare treatments with controls.
Since
informal trials will not be replicated from area to area,
but will each represent a complete experiment, ANOVA can
be used to test whether or not treated areas differ form
control areas. In most cases the treatment effects will
be random and quantitative, so that Duncan's multiple range
test can be used to test the significance of means within
treatments. Although such an approach will mean that results
from one area cannot be extrapolated to the site association
as a whole, the tests will provide meaningful results which
will be useful in directing future research.
The
formal and informal trials, and the monitoring areas described
in this Working Plan represent the first stage in a long
term, adaptive management approach. Information from all
three levels of trial will be useful in determining an optimal
approach to regenerating forest stands with high timber
and forage values. Therefore, the adaptive management approach
requires a long term commitment for all agencies involved
to achieve the objectives of the project.
Monitoring:
Results
will be forthcoming year-by-year as the work proceeds, and
the information acquired form these studies can be incorporated
into the trials as they continue to be established. In the
long term these observations will provide information to
assist in the design of experiments, especially the size
and orientation of gaps within the trials. The research
will also provide basic information on the autecology of
grizzly bear forage species that will assist in developing
ecologically-based approaches aimed at maintaining forage
abundance within managed areas.
- Retrospective
Research (This work should be carried out during the
berry production season (June-August)).
-
Assessing
Gap Characteristics
-
Assessment
of Forage Abundance
-
Light
Measurements
-
Data
Analysis
Time
Frame:
Integration
1993-1994
Year 2 - Implementation of trials and retrospective research
Current
Status:
Year
Three
A
range of activities were completed in 1994. Activities included
trial layout, updating of trial records, development of
a database, development of stocking tables for inclusion
in the Grizzly Stocking guidelines, and analysis of TASS
growth and yield projections.
Costs:
See
Table 2 for the breakdown. Costs are presented for only
those watersheds where it is known that visitation is required
in 1993-1994. There will undoubtedly be additional watersheds
which will be brought to our attention over the course of
the year, and no provision has been made for these.
Total
Projected Costs for 1993-1994 = $58,444.04
References:
Walters
and Hilborn. 1978. Ecological optimization and adaptive
management. Ann. Rev. Ecol. System. 9: 157-188.
McLennan,
D.S. and T. Johnson. 1993. (WORKING PLAN) An Adaptive
Management Approach For Integrating Grizzly Bear Habitat
Requirements and Silvicultural Practices in Coastal B.C.:
Working Plan. Oikos Ecological Consultants, Smithers,
B.C. for Hamilton, Wildlife Branch, MoE, Victoria.
McLennan,
D.S. and T. Johnson. 1993. (PROPOSAL) An Adaptive
Management Approach for Integrating Grizzly Bear Habitat
Requirements and Silvicultural Practices in Coastal B.C.
(1993-1994) Year 2 - Implementation of Trials
and Retrospective Research. Wildlife Branch, MoE,
Victoria, B.C.
McLennan,
D.S. and T. Johnson. 1994. (PROPOSAL) An Adaptive
Management Approach for Integrating Grizzly Bear Habitat
Requirements and Silvicultural Practices in Coastal B.C.
(1994-1995) Year 3 - Implementation of Trials.
Wildlife Branch, MoE, Victoria, B.C.
McLennan,
D.S. and T. Johnson. 1994. An Adaptive Management Approach
for Integrating Grizzly Bear Habitat Requirements and Silvicultural
Practices in Coastal B.C. (1993-1994) - Canopy Gap Literature
Review Retrospective Study Methodology. MoE, Wildlife
Branch, Victoria, B.C.
McLennan,
D.S. and T. Johnson. 1994. Guidelines for Integrating
Grizzly Bear Habitat and Silviculture in Coastal B.C. -
Stocking Standards. MoE, Wildlife Branch, Victoria,
B.C.
B.C.
Environment and Ministry of Forests. 1992. Interim
Guidelines for Integrating Coastal Grizzly Bear Habitat
and Silviculture in the Vancouver Forest Region. MoF
and MoE, Victoria, B.C.
Johnson.
1995. Re: Summary Report of Activities For 1994/95 Grizzly
Forage Trial Contract. MoE, Wildlife Branch, Victoria,
B.C.
Johnson,
T. 1995. Summary of Tass Projections As A Tool
to Finalize Grizzly Bear Stocking Guidelines. Ministry
of Environment, Wildlife Branch, Victoria, B.C.
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